W internecie jest tyle informacji na temat tego teropoda choÄby tutaj znajduje sie tyle informacji o niepewnych gatunkach i samym carcharodontosaurus saharicis, Ĺźe szok. JeĹli chcecie, mogÄ spokojnie zrobiÄ poprawÄ opisu. CytujÄ:
Carcharodontosaurus Stromer, 1931
C. saharicus (Deperet and Savornin, 1925) Stromer, 1931
= Megalosaurus saharicus Deperet and Savornin, 1925
= Dryptosaurus saharicus (Deperet and Savornin, 1928)
Albian (-Cenomanian?),
Continental Intercalaire, Algeria; (Baharija Formation, Egypt?; Kem Kem Formation, Morocco?)
Holotype- (destroyed) two teeth
Referred- ?(CMN 41817) tooth (54x27x12 mm) (Russell, 1996)
?(CMN 41818) tooth (67x36x22 mm) (Russell, 1996)
?(CMN 41819) tooth (69x34x16 mm) (Russell, 1996)
?(CMN 41859) maxillary fragment (Russell, 1996)
?(CMN 41908) tooth (30x24x11 mm) (Russell, 1996)
?(CMN 41910) tooth (23x19x6 mm) (Russell, 1996)
?(CMN 50792) cervical vertebra (148 mm) (Russell, 1996)
?(FPDM-V6211) fragmentary skull (Azuma, 2005)
?(IPHG 1912 VIII 68) ilium (Stromer, 1934)
(IPHG 1922 X46; holotype of Carcharodontosaurus) partial maxilla, maxillary teeth, nasals, frontals, parietals, supraoccipital, partial exoccipital-opisthotics, axis, anterior cervical vertebra (100 mm), cervical vertebra, proximal dorsal rib, proximal caudal vertebra (145 mm), proximal chevron (150 mm), partial chevron, manual ungual, incomplete pubes (>1 m), partial ischium, femora (1.26 m), fibula (880 mm) (Stromer, 1931)
?(MNNHN coll.) twelve teeth (to FABL of 42 mm), few mid caudal vertebrae (~70-100 mm) (Lapparent, 1960)
(SGM-Din 1) (12.79 m) incomplete skull (missing premaxillae, squamosals, quadratojugals) (~1.6 m) (Sereno et al., 1996)
? teeth, ilium (Lavocat, 1954)
? teeth (Taquet, 1976)
?teeth (Sadleir, 1998)
?remains (Smith et al., 2001)
Comments- Megalosaurus saharicus is based on two teeth from the Albian Continental Intercalaire of Algeria (Deperet and Sevornin, 1925). Later, Stromer (1931) described a partial skeleton from the Early Cenomanian Baharija Formation of Egypt and referred it to this species, creating the genus Carcharodontosaurus for it. An incomplete skull from the Cenomanian Kem Kem Formation of Morocco closely resembling Stromer's specimen was reported by Sereno et al. (1996). Brusatte and Sereno (2005) report a new species of Carcharodontosaurus, which differs in some cranial characters from Stromer's and Sereno et al.'s specimens. Dental differences are not reported, however, making referral of isolated teeth to either species uncertain. In fact, no diagnostic differences have been noted between the teeth of Carcharodontosaurus, Giganotosaurus and Mapusaurus. Thus the referral of isolated teeth to any particular derived carcharodontosaurid taxon is based solely on locality. This includes the holotype of Megalosaurus saharicus, perhaps indicating Stromer's specimen should be made a neotype for the species. The ilium (IPHG 1912 VIII 68) referred to Carcharodontosaurus by Stromer (1934) does not necessarily belong to this taxon, and may be abelisauroid (?= Deltadromeus or Bahariasaurus). Twelve teeth and a few mid caudal vertebrae from the type locality were described by Lapparent (1960). The teeth are described as having enamel wrinkles as in derived carcharodontosaurids and being very similar to the holotype, though the caudals' referral is uncertain. They are referred here to C. saharicus based on provenance. Russell (1996) described a maxillary fragment, cervical vertebra and teeth from the Kem Kem Formation of Morocco, which though closely resembling those in Stromer's specimen, are only referred to the species saharicus based on provenance. The same can be said for teeth from that locality mentioned by Lavocat (1954) and Sadlier (1998), though Lavocat's teeth were not only compared to Carcharodontosaurus, but also Tendaguru taxa and Tyrannosaurus, so may not be referrable to carcharodontosaurids at all. Smith et al. (2001) reported new remains of cf. Carcharodontosaurus from the Baharija Formation of Egypt. Sereno et al. (1996) referred the specimen of Spinosaurus B and the material of Sigilmassasaurus to Carcharodontosaurus saharicus, but this is not followed here (see Sigilmassasaurus entry).
References- Deperet and Savornin, 1925. Sur Ia decouverte d'une faune de vertebres albiens a Timintoun (Sahara occidental). C. R. Acad. Sci. Paris 181:1108-1111.
Stromer, 1931. Wirbeltiere-Reste der Baharijestufe (unterstes Cenoman). Ein Skellett-Rest von Carcharodontosaurus nov. gen.. Abh. Bayer. Akad. Wiss., Math.-Nat. Abt. (N. F.) 9 1-23, 1 pl.
Stromer, 1934. Ergebnisse der Forschungsreisen Prof. E. Stromers in den WĂźsten Ăgyptens. II. Wirbeltierreste der BaharĂŽje-Stufe (unterstes Cenoman). 13. Dinosauria. Abh. Bayer. Akad. Wiss., Math.-Nat. Abt., (n. s.) 22 1-79, 3 pls.
Lavocat, 1954. Sur les Dinosauriens du continental intercalaire des Kem-Kem de la Daoura. C. R. 19th Internatl. Geol. Congr. 1952: 65-68.
Lapparent, 1960. Les dinosauriens du "Continental intercalaire" du Sahara central. Mem. Soc. Geol. France 88A: 1-57.
Taquet, 1976. Geologie et Paleontologie du gisement de Gadoufaoua (Aptian du Niger): Cahires Paleont, 191pp.
Rauhut, 1995. Zur systematischen Stellung der afrikanischen Theropoden Carcharodontosaurus Stromer 1931 und Bahariasaurus Stromer 1934. Berliner geowissenschaftliche Abhandlungen E16 (Gundolf-Ernst-Festschrift): 357-375.
Russell, 1996. Isolated dinosaur bones from the Middle Cretaceous of the Tafilalt, Morocco. Bulletin du Muse'um national d'Histoire naturelle (4e se'r.) 18:349-402.
Sereno, Dutheil, Iarochene, Larsson, Lyon, Magwene, Sidor, Varricchio and Wilson, 1996. Predatory Dinosaurs from the Sahara and Late Cretaceous Faunal Differentiation. Science 272(5264): 986-991.
Sidleir, 1998. Theropod teeth from the Cretaceous of Morocco. JVP 18(3) 74A.
Larsson, 2001. Endocranial anatomy of Carcharodontosaurus saharicus (Theropoda: Allosauroidea) and its implications for theropod brain evolution. pp. 19-33. in Tanke, Darren H. & Carpenter, Kenneth, eds., 2001. Mesozoic Vertebrate Life: New Research inspired by the Paleontology of Philip J. Currie, Indiana University Press, Bloomington & Indianapolis, Indiana: xviii + 542 pp.
Smith, Lamanna, Lacovara, Dodson, Smith, Poole, Giegengack and Attia, 2001. A giant sauropod dinosaur from an Upper Cretaceous mangrove deposit in Egypt. Science 292:1704-1706.
Azuma, 2005. The Flying Dinosaurs: Fukui Prefectural Dinosaur Museum, 118pp.
Brusatte and Sereno, 2005. A new specis of Carcharodontosaurus (Dinosauria: Theropoda) from the Cenomanian of Niger and its implications for allosauroid phylogeny. JVP 25(3), 40A.
C. sp. nov. (Brusatte and Sereno, 2005)
Cenomanian, Late Cretaceous
Echkar Formation of the Tegama Group, Niger
Material- partial maxilla
Referred- braincase, teeth, anterior dentary
Diagnosis- (after Brusatte and Sereno, 2005) ventral margin of antorbital fossa not laterally swollen; robust medial ridge on maxilla; more slender, laterally divergent laterosphenoid alae; unspecified maxillary and endocast characters.
Comments- This is reportedly a new species of Carcharodontosaurus, with both Stromer's (1931) partial skeleton and Sereno et al.'s (1996) skull belonging to another species, provisionally called C. saharicus (though the holotype teeth may not be diagnostic to species level; see C. saharicus entry). However, Mapusaurus shares the laterally swollen ventral antorbital fossa margin with C. saharicus, but not C. sp. nov.. Furthermore, the synapomorphies uniting C. saharicus with C. sp. nov. to the exclusion of other carcharodontosaurids are as yet unreported.
References- Sereno, Wilson, and Conrad, 2004. New dinosaurs link southern landmasses in the mid-Cretaceous. Proceedings of the Royal Society of London B. 271(1546):1325-1330.
Brusatte and Sereno, 2005. A new specis of Carcharodontosaurus (Dinosauria: Theropoda) from the Cenomanian of Niger and its implications for allosauroid phylogeny. JVP 25(3), 40A.
C? sp. indet. (Lapparent, 1953)
Albian-Early Cenomanian, Early Cretaceous-Late Cretaceous
Tegama Group, Niger
Material- (MNNHN coll.; from In Abangarit) two braincase fragments, 137 teeth (anterior teeth- 80x33 mm, 77x31 mm, 62x28 mm, 64x27 mm, 54x26 mm; lateral teeth- 125x47 mm, 70x45 mm, 105x40 mm, 90x37 mm, 87x36 mm), proximal caudal vertebra (120 mm), distal caudal vertebra, manual phalanx II-2 (60 mm), pedal ungual II (90 mm) (Lapparent, 1960)
Comments- This material was not found associated, so could belong to multiple individuals of several large theropod taxa (Deltadromeus, Bahariasaurus, Spinosaurus, Sigilmassasaurus, etc.). Over a hundred of the teeth are carcharodontosaurid, however. Carcharodontosaurid material may be referrable to Carcharodontosaurus sp. nov., based on provenance.
References- Lapparent, 1953. Gisements de dinosauriens dans le "Continental intercalaire" d'In Abangharit (Saharia meridional): Compte rendu hebdomadaire des seances de l’Academie des Sciences Paris, v. 236, p. 1905-1906.
Lapparent, 1960. Les dinosauriens du "Continental intercalaire" du Sahara central. Mem. Soc. Geol. France 88A: 1-57.
C? sp. indet. (Schluter and Schwarzhans, 1978)
Albian, Early Cretaceous
Chenini Formation, Tunisia
Material- four teeth
References- Schluter and Schwarzhans, 1978.
Bouaziz, Buffetaut, Ghanmi, Jaeger, Martin, Mazin and Tong, 1988. Nouvelles decouvertes de vertebres fossiles dans l'Albien du Sud tunisien: Bulletin de la societie geologiques de France, 8th series, tomo 4, n. 2, p. 335-339.
Benton, Bouaziz, Buffetaut, Martill, Ouaja, Soussi and Trueman, 2000. Dinosaurs and other fossil vertebrates from fluvial deposits in the Lower Cretaceous of Southern Tunisia: Palaeogeography, Palaeoclimatology, Palaeoecology, v. 157, p. 227-246.
Buffetaut and Ouaja, 2002. A new specimen of Spinosaurus (Dinosauria, Theropoda) from the Lower Cretaceous of Tunisia, with remarks on the evolutionary history of the Spinosauridae: Bulletin de la societie geologiques de France, tomo 173, n. 5, p. 415-421.
C? sp. indet. (Lapparent, 1951)
Early Cretaceous
Dahar, Tunisia
Material- nine teeth
Reference- Lapparent, 1951. Decouverte de dinosauriens associes la und faunad de reptiles et de poissons, dans le Cretaceous inferieur de l'extreme sud tunisien: Compte rendu hebdomadaire des seances de l’Academie des Sciences Paris, v. 232, p. 1430-1432.
C? sp. indet. (Lapparent, 1960)
Albian-Early Cenomanian, Early Cretaceous-Late Cretaceous
Continental Intercalaire, Tunisia
Material- (MNNHN coll.) five teeth (Lapparent, 1960)
Comments- These teeth are said to have Carcharodontosaurus' 'characteristic thickness and form', so may be carcharodontosaurid.
Reference- Lapparent, 1960. Les dinosauriens du "Continental intercalaire" du Sahara central. Mem. Soc. Geol. France 88A: 1-57.
C? sp. indet. (Lapparent, 1960)
Aptian-Albian, Early Cretaceous
Elrhaz Formation, Niger
Material- (MNNHN Td. 2250) teeth (Taquet, 1976)
(MNNHN Td. 2269) teeth (Taquet, 1976)
(MNNHN coll.) anterior tooth (70 mm), lateral tooth fragment, mid caudal vertebra (>85 mm), radius (~110 mm), manual ungual (85 mm), pedal phalanx (85 mm) (Lapparent, 1960)
Comments- This material described by Lapparent was not found associated, so could belong to multiple individuals of several large theropod taxa (Deltadromeus, Bahariasaurus, Spinosaurus, Sigilmassasaurus, etc.). The tooth fragment has carcharodontosaurid enamel wrinkles.
References- Lapparent, 1960. Les dinosauriens du "Continental intercalaire" du Sahara central. Mem. Soc. Geol. France 88A: 1-57.
Taquet, 1976. Geologie et Paleontologie du gisement de Gadoufaoua (Aptian du Niger): Cahires Paleont, 191pp.
C? sp. indet. (Lapparent, 1960)
Berriasian-Barremian, Early Cretaceous
Irhazer Group, Niger
Material- (MNNHN coll.) partial cervical vertebra, two dorsal vertebrae (120 mm), two sacral vertebrae (280 mm combined), mid caudal vertebra (115 mm), three caudal vertebrae (100, 110, 120 mm), two distal chevrons (Lapparent, 1960)
Comments- This material was not found associated, so could belong to multiple individuals of several large theropod taxa (Deltadromeus, Bahariasaurus, Spinosaurus, Sigilmassasaurus, etc.).
Reference- Lapparent, 1960. Les dinosauriens du "Continental intercalaire" du Sahara central. Mem. Soc. Geol. France 88A: 1-57.
C? sp. indet. (Lapparent, 1960)
Early Cretaceous
Continental Intercalaire, Niger
Material- (MNNHN coll.; from Tefidet) two mid caudal vertebrae (85 mm) (Lapparent, 1960)
(from Akarazeras) teeth (Taquet, 1976)
Comments- This material was not found associated, so could belong to multiple individuals of several large theropod taxa (Deltadromeus, Bahariasaurus, Spinosaurus, Sigilmassasaurus, etc.).
References- Lapparent, 1960. Les dinosauriens du "Continental intercalaire" du Sahara central. Mem. Soc. Geol. France 88A: 1-57.
Taquet, 1976. Geologie et Paleontologie du gisement de Gadoufaoua (Aptian du Niger): Cahires Paleont, 191pp.
C? sp. indet. (Lapparent, 1960)
Early Cretaceous
Continental Intercalaire, Sahara Desert
Material- (MNNHN coll.) eight vertebrae, partial humerus, distal manual phalanx (Lapparent, 1960)
Comments- This material was not found associated, so could belong to multiple individuals of several large theropod taxa (Deltadromeus, Bahariasaurus, Spinosaurus, Sigilmassasaurus, etc.).
Reference- Lapparent, 1960. Les dinosauriens du "Continental intercalaire" du Sahara central. Mem. Soc. Geol. France 88A: 1-57.
C? sp. indet. (Buffetaut, 1989)
Late Albian-Early Cenomanian, Early Cretaceous
Tegama Formation?, Morocco
Reference- Buffetaut, 1989. New remains of the enigmatic dinosaur Spinosaurus from the Cretaceous of Morocco and the affinities between Spinosaurus and Baryonyx. Neues Jahrbuch fßr Geologie und Paläontologie Monatshefte 1989(2):79-87.
C? sp. indet. (Lapparent, 1960)
Albian, Early Cretaceous
Continental Intercalaire, Algeria
Material- ?(MNNHN coll.; from Alrar) caudal vertebra (60 mm), pedal ungual III (100 mm) (Lapparent, 1960)
(MNNHN coll.; from Aoulef) dorsal centrum (75 mm), proximal caudal centrum (105 mm), few mid caudal vertebrae (~70-100 mm), caudal vertebra (80 mm) (Lapparent, 1960)
(from Oued Boudjihane) teeth (Bassoullet and Iliou, 1967)
Comments- This material was not found associated, so could belong to multiple individuals of several large theropod taxa (Deltadromeus, Bahariasaurus, Spinosaurus, Sigilmassasaurus, etc.). A manual ungual from Dijoua referred to C. saharicus by Lapparent matches bone taxon J of Russell (1996), a possible oviraptorosaur. A distal metatarsal from Alrar referred to C. saharicus by Lapparent is actually a manual phalanx and matches bone taxon I of Russell, which may belong to the same species as bone taxon J.
References- Lapparent, 1960. Les dinosauriens du "Continental intercalaire" du Sahara central. Mem. Soc. Geol. France 88A: 1-57.
Bassoullet and Iliou, 1967. Discovery of dinosaurs associated with crocodilians and fish in the Lower Cretaceous of the Saharan Atlas (Algeria). SociĂŠtĂŠ GĂŠologique de la France, Comptes Rendus Sommaire des Sciences. 1967:294-295.
Russell, 1996. Isolated dinosaur bones from the Middle Cretaceous of the Tafilalt, Morocco. Bulletin du Muse'um national d'Histoire naturelle (4e se'r.) 18, 349-402.
C? sp. indet. (Bond and Bromley, 1970)
Late Jurassic or Early Cretaceous
Gokwe Formation, Zimbabwe
Material- teeth
Reference- Bond and Bromley, 1970. Sediments with the remains of Dinosaurs near Gokwe, Rhodesia: Palaeogeography, Palaeoclimatology, Palaeoecology, v. 8, p. 313-327.